Eumalacostraca is a subclass of crustaceans, containing almost all living malacostracans, or about
40,000 described species.[1] The remaining subclasses are the Phyllocarida and possibly
the Hoplocarida.[2] Eumalacostracans have 19 segments (5 cephalic, 8 thoracic and 6 abdominal).
This arrangement is known as the "caridoid facies", a term coined by William Thomas Calman in
1909. The thoracic limbs are jointed and used for swimming or walking. The common ancestor is
thought to have had a carapace, and most living species possess one, but it has been lost in some
subgroups.
Caridoid facies[edit]
Calman identified the following features as distinguishing eumalacostracan crustaceans: [3]
"Carapace enveloping the thoracic region; movably stalked eyes; biramous first antenna; scale-like
exopod on the second antenna; natatory exopods on the thoracic limbs; elongate, ventrally flexible
abdomen; tail fan formed by the lamellar rami of the uropods on either side of the telson."
Classification[edit]
Martin and Davis present the following classification of living eumalacostracans into orders, to which
extinct orders have been added, indicated by .[2]
The group as originally described by Karl Grobben[4] included the Stomatopoda (mantis shrimp), and
some modern experts continue to use this definition. This article follows Martin and Davis in
excluding them; they are placed in their own subclass, Hoplocarida.
Subclass Eumalacostraca Grobben, 1892
Superorder Syncarida Packard, 1885
Order Palaeocaridacea
Order Bathynellacea Chappuis, 1915
Order Anaspidacea Calman, 1904 (including Stygocaridacea)
Superorder Peracarida Calman, 1904
Order Spelaeogriphacea Gordon, 1957
Order Thermosbaenacea Monod, 1927
Order Lophogastrida Sars, 1870
Order Mysida Haworth, 1825
Order Mictacea Bowman, Garner, Hessler, Iliffe & Sanders, 1985
Order Amphipoda Latreille, 1816
Order Isopoda Latreille, 1817
Order Tanaidacea Dana, 1849
Order Cumacea Kryer, 1846
Superorder Eucarida Calman, 1904
Order Euphausiacea Dana, 1852
Order Amphionidacea Williamson, 1973
Order Decapoda Latreille, 1802