You've Evolved!: The Process of Evolution
You've Evolved!: The Process of Evolution
You’ve Evolved!
The Process of Evolution
Fossil Record
Fossils are the
preserved remains of
once-living organisms.
Fossils are created when
three (3) events occur:
1.) the organism must
become buried in
sediment; 2.) the
calcium in the bone or
other hard tissue must
mineralize; and 3.) the
surrounding sediment http://dailymail.co.u
must eventually harden Source: http://dailymail.co.uk
to form rock. The process of fossilization probably occurs rarely.
Usually, animals or plant remains will decay or be scavenged
before the process can begin. Only a fraction of the species that
have existed are known from fossils. Nonetheless, the fossils
that have been discovered are sufficient to provide detailed
information on the course of evolution through time.
th
Source: Raven, et al., Biology 9 edition figure 21.10
Dating Fossils
By dating the rocks in which fossils occur, we can get an Today, rocks are dated by measuring the degree of decay of
accurate idea of how old fossils are. In Darwin’s day, rocks were certain radioisotopes contained in the rock, a process called
dated by their position with respect to one another, a process absolute dating; the older the rock, the more of its isotopes have
called relative dating; rocks in deeper strata are generally older. decayed. Because radioactive isotopes decay at a constant rate
unaltered by temperature or pressure, the isotopes in a rock act to evolutionary modification of the limbs. All three (3) fossil
as an internal clock, measuring the time since the rock was forms occurred in the Eocene Period, 45-55 million years ago
formed. This is a more accurate way of dating rocks and (MYA).
provides dates stated in millions of years, rather than relative
dates. When fossils are arranged according to their age, from Modern-day members of the Equidae include horses,
oldest to youngest, they often provide evidence of successive zebras, and donkeys, all of which are large, long-legged, fast-
evolutionary change. At the largest scale, the fossil record running animals adapted to living on open grasslands. These
documents the progression of life through time, from the origin species, all classified in the genus Equus, are the last living
of eukaryotic organisms, through the evolution of fishes, the rise descendants of a long lineage that has produced 34 genera since
of land-living organisms, the reign of dinosaurs, and on to the its origin on the Eocene Period approximately 55 MYA.
origin of humans. Examination of these fossils provided a particularly well-
documented case of how evolution has proceeded by adaptation
Given the low likelihood of fossil preservation and to changing environments.
recovery, it is not surprising that there are gaps or “missing
links” in the fossil record. Nonetheless, paleontologists
(scientists who study fossils) continue to fill the gaps in fossil
record. Today, fossil record is far more complete, particularly
among vertebrates; fossils have been found linking all the major
groups. For example, a four-legged aquatic mammal was
discovered that provides important insights concerning the
evolution of whales and dolphins from land-living hoofed
ancestors. Similarly, a fossil snake with legs has shed light on
evolution of snakes, which descended from lizards that
gradually became more and more elongated with simultaneous
reduction and eventual disappearance of limbs.
The earliest known members of the horse family are species All these changes may be understood as adaptation to
in the genus Hyracotherium, which did not look much like changing global climates. In particular, during the late Miocene
horses at all. Small, with short legs and broad feet, these species and early Oligocene (20-25 MYA), grasslands became
occurred in wooded habitats, where they probably browsed on widespread in North America, where much of horse evolution
leaves and herbs and escaped predators by dodging through occurred. As horses adapted to these habitats, long-distance and
openings in the forest vegetation. The evolutionary path from high speed locomotion probably became more important to
these small creatures to the workhorses of today has involved escape predators and travel great distances.
changes in a variety of traits, including: By contrast, the greater flexibility provided by multiple
Size – the first horses were no bigger than dogs, some toes and shorter limbs, which was advantageous for ducking
was the size of a housecat. The modern equids can weigh through complex vegetation, was not longer beneficial. At the
more than half a ton. same time, horses were eating grasses and other vegetation that
Toe reduction – the first horses had four (4) toes on its contained grit and other hard substances, thus favoring teeth and
front feet and three (3) on its hind feet; encased in fleshy skulls better suited for withstanding such materials.
pads. The feet of modern horses have a single toe
enclosed in a tough bony hoof. Homology
Tooth size and shape – the teeth of Hyracotherium were As vertebrates evolved, the same bones were sometimes put to
small and relatively simple in shape. Through time, horse different uses. For example, the forelimbs of vertebrates are
teeth have increased greatly in length and have homologous structures, that is, structures with different
developed a complex pattern of ridges on their molars appearances and functions that are all derived from the same
and premolars. Accompanying these changes have been body part in a common ancestor.
alterations in the shape of the skull that strengthen the
skull to withstand the stresses imposed by continual
chewing.
Embryological Development embryological forms provides convincing evidence that they are
both mollusks
In many cases, the evolutionary history of an organism can
be seen to unfold during its development, with the embryo Vestigial Structures
Many organisms
possess vestigial
structures that
have no apparent
function, but
resemble
structures their Source: Raven, et al., Biology 5th edition figure
21.17
presumed
ancestors had. Human, for example, possess a complete set of
muscles for wiggling their ears, just as a coyote does. Boa
Source: Raven, et al., Biology 5th edition figure 21.15 constrictors have hip bones and rudimentary legs. The skeleton
of a baleen whale, which contains pelvic bones, as other
exhibiting characteristics of the embryos of its ancestors. For mammal skeletons do, even though such bones serve no known
example, early in their development, human embryos function in the whale. It is difficult to understand vestigial
possess gill slits, like fish; at later stage, every human structures such as these as anything other than evolutionary
embryo has a long bony tail, the vestige of which we carry relics, holdovers from the evolutionary past. They argue strongly
to adulthood as the coccyx at the end of our spine. Human for the common ancestry of the members of the groups that share
fetuses even possess a fine fur (called lanugo) during the them, regardless of how different they have subsequently
fifth month of development. These relict developmental become.
forms suggest that our development has evolved with new
instructions layered on top. Evidence from Biochemistry and Genetics
The observation that seemingly different organisms may Blood tests show that human blood is more like the
exhibit similar embryological forms provides indirect but rabbit’s blood than a frog’s blood or an insect’s blood.
convincingly evidence of a past evolutionary relationship. Slugs Furthermore, human blood is more like an ape blood is more like
and giant ocean squids, for example do not bear much superficial an ape’s blood than rabbit’s blood. Notice that the closer the
resemblance to each other, but the similarity of their groups are in structure, the closer their blood chemistry. This is
one more piece of evidence of relationship and common As a result, the population will gradually come to
ancestral origin. include more and more individuals with advantageous
characteristics. In this way, the population evolves and
Modern genetics also provides evidence of evolution. becomes better adapted to its local circumstances.
All organisms use the same genetic code to synthesize proteins.
A universal genetic code is consistent with the idea that all THREE (3) SAMPLES OF NATURAL SELECTION
organisms evolved from a common ancestor that used that code.
Selection to Avoid Predators
Theories of Evolution
Selection
Natural Selection In these areas, the black rock formations produced when
lava cooled contrasts to the surrounding bright glare of the
Natural selection produces evolutionary change dessert sand. The pocket mice in figure (A) matches the color of
when in a population, some individuals which possess the cooled lava and the one of figure (B) is light colored to match
certain inherited characteristics, produce more surviving the white sand.
offspring than individuals lacking these characteristics.
Selection to Match Climatic Conditions uptake of insecticide, whereas alleles at the kdr and dld-r genes
decrease the number of target sites, thus decreasing the binding
Many studies of ability of insecticide. Other alleles enhance the ability of the
selection have focused on insects’ enzymes to identify and detoxify insecticide molecules.
genes encoding enzymes.
Often investigators fined the
enzyme allele frequencies Source: http://new.coolclassrooom.org
vary with latitude, so that one
allele is more common in northern populations, but is
progressively less common at more southern locations. An
example is seen in studies of a fish, the mummichog (Fundulus
heteroclitus), which ranges along the eastern coast of North
America. In this fish, geographic variation occurs in allele
frequencies for the gene that produces the enzyme lactate
dehydrogenase, which catalyzes the conversion of pyruvate to
lactate (for release of energy when there is not enough oxygen). th
Source: Raven, et al., Biology 9 edition figure 20.7
Biochemical studies show that enzymes formed by these alleles
function differently at different temperatures, thus explaining
their geographic distributions. The form of the enzyme more
frequent in the north is a better catalyst at low temperatures than
the enzyme from the south. Moreover, studies indicate that at
low temperatures, individuals with the northern allele swim
faster, and presumably survive better, than individuals with the
alternative alleles.
The Relationship and Differences of Natural Selection and suggested to him that lineages change gradually as species
Evolution migrate from one area to another.
Natural selection is a process, whereas evolution is the On the Galapagos Islands, Darwin encountered a variety
historical record, or outcome, of change through time. An of different finches on various islands. The 13 species of finches,
example is how giraffes evolved to have longer necks from their although related slightly differed in appearance, particularly in
ancestors. Through the process of natural selection, longer- their beaks. Darwin thought that all these birds had descended
necked giraffes are more successful in survival and pass the from a common ancestor from the South American mainland
long-neck trait on to their offspring. several million years ago. Eating different foods on different
islands, the finches' beaks had changed during their descent.
Natural selection can lead to evolution, but natural
selection is only one of several processes that can result to The diversity of Darwin’s finches:
evolutionary change. The five agents of evolutionary change are
mutation, gene flow, nonrandom mating, and genetic drift. Ground finches – feed on seeds that they crush in their
powerful beaks
Natural selection can occur without producing
evolutionary change; only if variation is genetically based will Warbler finches – feed on insects with their smaller and
natural selection lead to evolution. An example is an offspring narrower beaks
who do not resemble its parents because of its experience on
different environments during growth and not because it is Fruit/bud/cactus finches – feed on fruit/bud/cactus and
different genetically. other insects they attract
Charles Darwin is an English Naturalist who wrote the Sharp-beaked ground finches – include “vampires” that
book “On the Origin of Species by Means of Natural Selection”. sometimes creep up on seabirds and use their sharp beaks
In Darwin’s time, most people believed that the different kinds to pierce seabirds’ skin and drink their blood.
of organisms resulted from direct actions of a creator. Darwin
was a part of a five-year navigational mapping expedition Wood-pecker finches – pick up a twig, cactus spine, or
around the coast of South America, aboard H.M.S. Beagle. leaf stalk, and trim it into shape with their beaks, and
During the voyage, he studied a wide variety of plants and then poke it into dead branches to pry out grubs.
animals on continents and islands in distant seas. Darwin
repeatedly saw characteristics of similar species varied
somewhat from place to place. These geographical patterns
As basis, for his observations, these assumptions were strong running muscles in its legs. An animal like the giraffe has
developed: to keep reaching up to tall shrubs and trees for food. Each
1. The Theory of Need generation of giraffes had a slightly longer neck. After hundreds
2. The Theory of Use and Disuse of thousands of years, the short-necked giraffes became the
3. The Theory of Transmitting Acquired Traits present-day long necked giraffes. In the case of snakes, the legs
and short bodies, which were believed to be present in their early
Theory of Need ancestors, eventually disappeared. It became necessary for
Lamarck believed that organisms change because they snakes to walk through narrow places. They began to stretch
need to. It seemed, according to him, that there is natural their bodies and to crawl rather than walk.
tendency for organisms to make themselves better and more
suited to their environment. Theory of Acquired Traits
For example, birds developed wings because their Lamarck explained that an organism acquired a body
ancestors developed the need to fly. Elephants have long trunks structure (for example, characteristic structure to its offspring.
for gathering food; lions have powerful claws and jaws for He thought that if an organism acquired a body structure
capturing prey; and deer have long, swift legs for escaping (for example, long limbs, legs, or necks) during its lifetime, it
predators. Each organism therefore adapts to suit its particular could pass on that characteristic structure to its offspring.
way of life. According to him, this is how evolution occurs.
Lamarck further believed that changes happened
because organisms were able to reshape their bodies in many Lamarck and Darwin’s Observations Compared
new ways. Body structures, he explained, could be altered in size
or shape depending on the needs of the organism. Lamarck’s giraffe
For example, birds trying to use their front limbs for 1. Original, short-necked ancestor
flying could eventually change those limbs into wings, the same 2. Keeps stretching neck to reach leaves higher up on tree
way birds with short legs could develop long legs by stretching 3. And stretching
into deeper water. Then, if these animals had no more need for 4. And stretching until neck becomes progressively longer
the structures they had developed through time, these body parts
would decrease in size or eventually disappear.
this handout for the discussion of the two (2) types of selection Anthropoids
which are natural and artificial. Known as the higher primates; include monkeys,
apes, and humans.
The Evolution of Humans Diurnal, feeding mainly in fruits and leaves.
Live in groups of complex interactions.
In 1897, Charles Darwin published another ground Tend to care for their young for prolonged periods,
breaking book, “The Descent of Man”. In this book, he allowing for a long childhood of learning and brain
suggested that humans evolved from the same African ape development.
ancestors that gave rise to the gorilla and the chimpanzee. Little New World monkeys - anthropoids migrated to
fossil evidence existed at that time to support Darwin's case but South America
numerous fossil discoveries made since then strongly support his Old World monkeys – a group of anthropoids that
hypothesis. remained in Africa.
The story of human evolution begins around 65 million
years ago, with a group of small, arboreal (species that live in Homonoids
trees) mammals called Archonta. These primarily insectivorous
Apes have larger brains than monkeys and they lack
mammals had large eyes that were most likely nocturnal (active tails. The living apes consists of the gibbon,
at night). Their radiation gave rise to different types of orangutan, gorilla, and chimpanzee. Fossils
mammals, including bats, tree shrews, and other primates, the discovered suggest the first possible Hominoid:
order of mammals that contains humans. o Ramapithecus – 8 million year old jaw with
teeth found in India. Fossils indicate they are
40 million years ago, the Archonta split in to two (2) groups: closely related to orangutan.
o Proconsul – has many characteristics of Old
Prosimians World monkeys, but lacks tail and has
Looked like a cross between a squirrel and a cat. apelike hands, feet, and pelvis.
Common in North America, Europe, Asia, and Apes evolved knuckle-walking, supporting their
Africa. weight on the back sides of their fingers.
Nocturnal, have grasping digits, and binocular Hominids became bipedal and walking upright.
vision.
An example of a prosimian that survived today is the Five to ten million years ago, the world’s climate began
tarsier. to get cooler, and the great forests of Africa were largely
replaced with savannas and open woodland. In response to Australopithecus boisei – nicknamed the “Nut-
these changes, the hominids, which were bipedal arose. cracker man”. It had a great bony ridge, a Mohawk
In every case where fossils allow determination to be haircut of bone, on the head to anchor its immense
made, the hominids are bipedal, walking upright. Bipedal jaw muscles.
locomotion is the hallmark of hominid evolution. Australopithecus afarensis - nicknamed “Lucy”. The
There are two (2) major groups of hominids: The skeleton was 40% complete and over 3 million years
Australopithecus and the Homo: old. The shape of the pelvis indicated that Lucy was
a female, and her leg bones proved she walked
Australopithecus upright. Her brain was no larger than that of a
First hominid chimpanzee, about the size of an orange.
Walked upright and lived in Africa over 3 million Homo
years ago African origin
Had a small brain Have larger brain than Australopithecus and most
Weighed about 18 kilograms and were about 1 meter likely used tools
tall Homo habilis – means “handy man”. Found with
Dentition was distinctly hominid, but their brains stone tools near its fossil. Its skull suggests a brain
were not any larger than those of apes volume of 680 cubic centimeters that which is larger
Australopithecus africanus – a skull of a five-year- than the Australopithecus. Homo habilis was small
old individual, still with his milk teeth. While the in stature, with arms longer than legs and a skeleton
skull had many apelike features such as a projecting much like Australopithecus.
face and a small brain, it had distinctly human Homo rudolfensis – the skull which was 1.9 million
features as well (a rounded jaw, unlike the pointed years old, had a brain volume of 750 cubic
jaw of apes). The ventral position of the foramen centimeters and many characteristics if human
magnum (the hole at the base of the skull from which skulls. It was assigned to a different species than
the spinal cord emerges) suggested that the creature Homo habilis because of its substantial brain
walked upright. Raymond Dart, the anatomy expansion.
professor who discovered the skull, concluded it was Homo ergaster – have even larger brains than Homo
a human ancestor. Scientists now estimate the skull rudolfensis, with skeletons less much like the
to be 2.8 million years old. Australopithecus and more like a modern human in
Australopithecus robustus – second kind of both size and proportion.
Australopithecus unearthed in South Africa. It had
massive teeth and bones.
09 Handout 1 *Property of STI
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SH1631